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| 1 | This article contains information about the complex insects that form and inhabit leaf mines in the forest-steppe of European Russia (Middle Volga). Mines of moths of the genus Phyllonorycter (Lepidoptera, Gracillariidae) described as an example. Hemiptera, Thysanoptera, Coleoptera, Hymenoptera and Trombidiformes associates with leaf mines these moths. Keywords: leaf mines, leaf-mining moths, parasitic wasps, eulophids, host plants, entomofauna, invasive species | 878 | ||||